科研之友
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摘要
The Apatesieae (Aizoaceae) is a small tribe of 11 species in seven genera, Apatesia, Carpanthea, Caryotophora, Conicosia, Hymenogyne, Saphesia and Skiatophytum. All are succulent annual or perennial herbs with mostly flat, rarely trigonous to terete leaves. Analysis of molecular data from four plastid and nuclear ITS sequences confirm that the two informal groups identified earlier within the Apatesieae, the "Apatesia-group" and the "Skiatophytum-group", as well as all genera were found to be monophyletic. The relationships among genera of the "Apatesia-group", however, differ from those previously suggested, which were largely based on characters of the fruits. Here we show that characters of the fruit are highly homoplasious. Our analysis of distributional data for the three major clades of the Aizoaceae with mesomorphic leaves, namely Apatesieae (Ruschioideae), Aizooideae and Mesembryanthemoideae, shows that each has different centres of diversity. The Apatesieae are most diverse in the south-western Cape, in the fynbos of the Cape Floristic Region (CFR) and they are restricted to the Greater Cape Floristic Region (GCFR). We revise the classification for the Apatesieae and include the monotypic genera Caryotophora and Saphesia in Skiatophytum, in order to increase the phylogenetic information conveyed by the classification. We show that the lectotype and protologue of Mesembryanthemum flaccidum do not correspond to the species currently associated with this name, which we therefore describe as a new species, i.e., Skiatophytum flaccidifolium. In addition, we describe a new subspecies, Hymenogyne glabra subsp. namaquensis, which is endemic to the Kamieskroon area in Namaqualand. Amended descriptions for Skiatophytum and Hymenogyne as well as an improved key to the genera of the Apatesieae and to the species of Skiatophytum are provided.
- 出版日期2015-6
