摘要

Prokop, J., Roques, P. & Nel, A., 2014. New non-holometabolous insects from the Pennsylvanian of the Avion locality in Pas-de-Calais, France (Insecta: Exopterygota'). Alcheringa 38, 155-169. ISSN 0311-5518.New palaeopteran and neopteran insects are described from Moscovian impression fossils of the Pas-de-Calais Basin in northern France. Spilaptera splendens sp. nov. (Palaeodictyoptera) is based on new forewing venation characters including a well-developed anal bar in the form of a cuticular reinforcement extending from the anal area to the radial stem, a feature previously unknown in this genus. This character was also confirmed after re-examination of S. libelluloides. The discovery of a wing part with a large anal fan supports the presence of a gigantic palaeodictyopteran that can be assigned to Homoiopteridae. Based on a petiolate wing, Jacquesoudardia magnifica gen. et sp. nov. represents a small damselfly-like odonatopteran that was probably predatory on small arthropods and coexisted with larger meganeurid species such as Gallotupus oudardi from the same locality. The discovery of Paolia sp. (Paoliidae), known previously from the Bashkirian of Indiana (USA) and South Limbourg (the Netherlands), extends the distribution and stratigraphic range of the genus. Characterized by a short ScP, Aviogramma gracilis gen. et sp. nov. is the second member of Caloneurodea from the Pas-de-Calais Basin after Pruvostiella lecomtei (Pruvost). Aviocladus pectinatus gen. et sp. nov. and Aviohapaloptera bethouxi gen. et sp. nov. are both assigned to Archaeorthoptera. Aviocladus pectinatus, a member of Cnemidolestodea, preserves a venation pattern that allows restoration of Tococladus to Archaeorthoptera. Aviohapaloptera has strongly similar venation to Hapaloptera, known from Mazon Creek (USA), in bearing a pectinate CuPb, CuA+CuPa with few posterior branches, a narrow costal area and radial posterior (RP) with few branches. In addition, we restore Narkeminidae stat. rest. and separate it from Proedischiidae. These new fossils demonstrate that the insect fauna from the Pas-de-Calais Basin was highly diverse with links to Bashkirian/Moscovian assemblages in Euramerica, such as the Mazon Creek (Illinois, USA) entomofauna.Jakub Prokop [jprokop@natur.cuni.cz], Charles University in Prague, Faculty of Science, Department of Zoology, Vinina 7, CZ-128 44, Praha 2, Czech Republic. Patrick Roques [patrick.roques93@wanadoo.fr] allee des Myosotis, Neuilly sur Marne, F-93330, France. Andre Nel [anel@mnhn.fr] CNRS UMR 7205, Museum National d'Histoire Naturelle, CP 50, Entomologie, 45 rue Buffon, F-75005 Paris, France. Received 15.7.2013, revised 10.9.2013, accepted 23.9.2013. ????Pas-de-Calais???Moscovian????????????????????? Spilaptera spledens sp. nov. (Palaeodictyoptera)???????????, ??????????????????????????????, ??????????, ?????S. libelluloides???????????????????????????????? ?????????Homoiopteridae??????, Jacquesoudardia magnifica gen. et sp. nov.????????????, ?????????, ????meganeurid???Gallotupus oudardi ??????? ? Paolia sp. (Paoliidae)?????????????????, ?????????????????(??)?????(??)????ScP??, Aviogramma gracilis gen. et sp. nov.?Pas-de-Calais???Pruvostiella lecomtei (Pruvost)???Caloneurodea??????? Aviocladus pectinatus gen. et sp. nov ?Aviohapaloptera bethoux gen. et sp. nov ???Archaeorthoptera ? Cnemidolestodea?????, Aviocladus pectinatus, ???????, ?????Tococladus????Archaeorthoptera ?Mazon Creek(??)? Aviohapaloptera???Hapaloptera??????, ????CuPb???????CuA +CuPa?????????????????(RP)???, ?????Narkeminidae stat. rest. ,???Proedischiidae??????????, ?Pas-de-Calais???????????????, ???Bashkirian/Moscovian???Mazon Creek(???????) ??????????

  • 出版日期2014-4-3