摘要

Based on a new, centennial-scale-resolution pollen record from the terrestrial site of Tenaghi Philippon (NE Greece) we report a series of high-amplitude millennial-scale forest expansion events during the interval from 312 to 240 ka (equivalent to isotopic substages c-a of Marine Isotope Stage [MIS] 9, MIS 8, and the onset of MIS 7e). Pollen-based indices support the climatic interpretation of the forest expansion events as indicators of abrupt increases in temperature and moisture availability in the Mediterranean region. Long-term changes in vegetation cover and composition during the studied interval permit the definition of glacial substages (early, middle and late) analogous to the glacial substages of the last climatic cycle. Within this framework, forest expansion events occurred during the early glacial (equivalent to MIS 9c-a) and the later part of the middle glacial (mid to late MIS 8), but are absent from the early part of the middle glacial (early MIS 8). Forest expansion events are also detected during the Lateglacial and onset of MIS 7, associated with a millennial-scale climate oscillation across the glacial-interglacial transition, i.e. Termination III (T-III). A peak-for-peak match between forest expansion events and Antarctic temperature maxima suggests interhemispheric coupling of millennial-scale climate variability via global oceanic circulation change and synergistic variability in atmospheric circulation consistent with the bipolar see-saw. The resolution of our record (mean: 290 yr) also permits the characterisation of climate evolution over the course of interstadial episodes following forest expansion events, and highlights differences in the characteristics of early and middle glacial interstadials (saw-tooth vs. plateau profile). These differences point to an influence of long-term boundary conditions, especially global ice volume, on the characteristics of millennial-scale events.